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Ceratopsians is a group of herbivorous, beaked dinosaurs
which thrived in what are now North America and Asia, during
the Cretaceous Period, although ancestral forms lived earlier,
in the Jurassic. Early members such as Psittacosaurus
were small and bipedal. Later members, including ceratopsids
like Centrosaurus and Triceratops, became
very large quadrupeds and developed elaborate facial horns and
a neck frill. While the frill might have served to protect the
vulnerable neck from predators, it may also have been used for
display, thermoregulation, the attachment of large neck and
chewing muscles or some combination of the above. Ceratopsians
ranged in size from 1 meter (3 ft) and 23 kilograms (50 lb)
to over 9 meters (30 ft) and 5,400 kg (12,000 lb).
Pachycephalosauria ("Thick headed lizards")
is a clade of ornithischian dinosaurs. Well-known genera include
Pachycephalosaurus, Stegoceras, Stygimoloch,
and Dracorex. Most lived during the Late Cretaceous
Period, in what is now North America and Asia. They were all
bipedal, herbivorous/omnivorous animals with thick skulls. In
some species the skull roof is domed and several inches thick;
in others it is flat or wedge-shaped. The dome may be surrounded
by nodes (for instance, Pachycephalosaurus), spikes
(Stygimoloch), or both (Dracorex).
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Achelousaurus |
Achelousaurus ("Achelous's lizard")
is a genus of centrosaurine ceratopsid dinosaur from the Late
Cretaceous Period of what is now North America. It was a quadrupedal
herbivore with a parrot-like beak, a rough boss (raised bony area)
on the snout and two more behind the eyes, and two horns on the
end of its long bony neck frill. With a total body length of 6
meters (20 feet), Achelousaurus was a medium-sized ceratopsian.
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Albertaceratops |
Albertaceratops ("Alberta horned
face") was a genus of centrosaurine horned dinosaur from
the middle Campanian-age Upper Cretaceous Oldman Formation of
Alberta, Canada, and Judith River Formation of Montana, USA. It
is known from a single complete skull found in August 2001 and
skull and postcranial fragments. This genus is unusual in combining
long brow horns with an otherwise centrosaurine skull, as centrosaurines
normally possess short brow horns. Over its nose was a bony ridge,
and on its frill were two large outwardly-projecting hooks. A
phylogenetic analysis carried out by its describer, Michael J.
Ryan, found it to be the most basal centrosaurine.
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Anchiceratops |
Anchiceratops ("Near horned face")
is a genus of chasmosaurine ceratopsid dinosaur from the Late
Cretaceous Period of western North America. Like other ceratopsids,
it was a quadrupedal herbivore with three horns on its face, a
parrot-like beak, and a long frill extending from the back of
its head. The two horns above the eyes were longer than the single
horn on its snout, as in other chasmosaurines. Anchiceratops
approached 20 feet (6 m) in length.
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Centrosaurus  |
Centrosaurus ("Pointed lizard")
is an herbivorous ceratopsian dinosaur from the late Cretaceous
of North America, approximately 75 million years ago. Centrosaurus
massive body was borne by stocky limbs, although at 18-20 ft (6m)
it was not a particularly large dinosaur. Like other centrosaurines,
Centrosaurus had a single large horn over the nose. It
may curve forwards or backwards in different species. A pair of
small horns is also found over the eyes while the frill of Centrosaurus
was moderately long, with fairly large fenestrae and small hornlets
along the outer edge.
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Chasmosaurus |
Chasmosaurus is a genus of ceratopsid
dinosaur from the Upper Cretaceous Period of North America. Its
name means "opening lizard", referring to the large
openings (fenestrae) in its frill. With a length of 5–6
metres (16–20 ft) and a weight of 3.6 tonnes (4.0 short
tons), Chasmosaurus was a ceratopsian of average size.
Like all ceratopsians, it was purely herbivorous. It was initially
to be called Protorosaurus, but this name had been previously
published for another animal.
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Einiosaurus  |
Einiosaurus is a medium-sized centrosaurine
("Short-frilled") ceratopsian from the Upper Cretaceous
(Campanian) Two Medicine Formation of northwestern Montana. Einiosaurus
is typically portrayed with a low, strongly forward-curving nasal
horn that resembles a bottle opener, though this may only occur
in some adults. Supraorbital (over-the-eye) horns are low and
rounded if present at all, as opposed to ceratopsids with prominent
supraorbital horns such as Triceratops. A pair of large
spikes projects backwards from the relatively small frill.
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Homalocephale 
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Homalocephale ("Even head")
is a genus of dinosaur belonging to the pachycephalosaurid family,
which lived during the late Cretaceous period. The genus was described
in 1974 by Osmólska and Maryañska, and consists
of a single species, the 3 metres (10 ft) long herbivore. Sporting
a flat, wedge-shaped skull roof, H. calathocercos was
different from other pachycephalosaurs. Nonetheless, the surface
of the skull was greatly thickened, and it had been proposed that
males competed in head-butting contests, similar to those of modern
iguanas. The species is also noted for having an unusually broad
pelvis, which lead some paleontologists to suggest that the wide
hips were for giving birth to live young. Others have suggested
that the width served to protect vital organs from harm during
flank-butting. Homalocephale also had rather long legs,
indicating a fast-moving gait.
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Pachycephalosaurus |
Pachycephalosaurus ("Thick headed
lizard") is a genus of pachycephalosaurid dinosaur. It lived
during the Late Cretaceous Period (Maastrichtian stage) of what
is now North America. Remains have been excavated in Montana,
South Dakota, and Wyoming. It was an herbivorous or omnivorous
creature which is only known from a single skull and a few extremely
thick skull roofs.
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Pachyrhinosaurus |
Pachyrhinosaurus ("Thick-nosed
reptile") is a genus of ceratopsid dinosaur from the Late
Cretaceous period of North America. Instead of horns, the skull
bears massive, flattened bosses, the largest being over the nose.
These were probably used in butting and shoving matches, as in
musk oxen. A single pair of horns grew from the frill and extended
upwards. It appears that that both the shape and size of the frill
was highly individualized, reliant on gender and perhaps other
factors. Pachyrhinosaurus is most closely related to
Achelousaurus.
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Pentaceratops |
Pentaceratops is a genus of ceratopsid
dinosaur from the late Cretaceous Period of what is now North
America. Its name means "five-horned face", in reference
to its two long epijugal bones, spikes which protrude out sidewards
from under its eyes, in addition to the three more obvious horns.
Pentaceratops lived around 75-73 million years ago and measured
about 8 m (27 ft) long, and has been estimated to have weighed
around 5,500 kg (13,000 lb).
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Prenocephale |
Prenocephale was a small pachycephalosaurid
dinosaur genus from the Late Cretaceous (from the Campanian through
to the Maastrichtian) and was similar in many ways to its close
relative, Homalocephale. Prenocephale probably
weighed around 130 kilograms (290 lb) and was around 2.4 metres
(8 ft) long. Unlike the flattened wedge-shaped skull of Homalocephale,
the head of Prenocephale was rounded and sloping. The
dome had a row of small, bony spikes and bumps. It lived in what
is now Mongolia; but in high upland forests, not the dry deserts
of Mongolia today.
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Protoceratops |
Protoceratops ("First Horned
Face") is a genus of sheep-sized (1.5 to 2 m long) herbivorous
ceratopsian dinosaur, from the Upper Cretaceous Period (Campanian
stage) of what is now Mongolia. It was a member of the Protoceratopsidae,
a group of early horned dinosaurs. Unlike later ceratopsians,
however, it lacked well-developed horns and retained some primitive
traits not seen in later genera.
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Psittacosaurus |
Psittacosaurus ("'Parrot lizard")
is a genus of psittacosaurid ceratopsian dinosaur from the Early
Cretaceous Period of what is now Asia, about 130 to 100 million
years ago. It is notable for being the most species-rich dinosaur
genus. At least ten extinct species are recognized from fossils
found in different regions of modern-day China, Mongolia and Russia,
with a possible additional species from Thailand.
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Stegoceras |
Stegoceras ("Horned roof")
was a genus of plant-eating ornithischian pachycephalosaurid dinosaur
that lived in what is now North America during the Late Cretaceous
period. It had an estimated length of up to 2 metres (6.6 ft).
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Stygimoloch  |
Stygimoloch ("Horned devil from
the river of death") is a genus of pachycephalosaurid dinosaur
from the end of the Cretaceous period, roughly 65 million years
ago. It is currently known from the Hell Creek Formation and Lance
Formation of the Western Interior (United States), where it lived
alongside Tyrannosaurus and Triceratops. It
is a relatively large pachycephalosaur, with the skull being about
46 centimeters long (18 in). Among North American pachycephalosaurs,
only Pachycephalosaurus is larger. Unlike other pachycephalosaurs,
the domed skull is relatively small, slightly flattened from side
to side, and pear-shaped; even when isolated this unusual dome
can easily be distinguished from the broader, larger domes of
Pachycephalosaurus.
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Styracosaurus |
Styracosaurus ("Spiked lizard"')
was a genus of herbivorous ceratopsian dinosaur from the Cretaceous
Period (Campanian stage), about 76.5 to 75.0 million years ago.
It had four to six long horns extending from its neck frill, a
smaller horn on each of its cheeks, and a single horn protruding
from its nose, which may have reached dimensions of around 60
centimeters (2 ft) long and 15 centimeters (6 in) wide. The function
or functions of the horns and frills have been the subject of
debate for many years. Styracosaurus was a large dinosaur,
reaching lengths of 5.5 meters (18 ft) and weighing nearly 3 tons.
It stood about 1.8 meters (6 ft) tall. Styracosaurus
possessed four short legs and a bulky body. Its tail was rather
short. It also had a beak and flat cheek teeth, indicating that
its diet was herbivorous. Like other ceratopsians, this dinosaur
may have been a herd animal, traveling in large groups, as suggested
by bonebeds.
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Torosaurus |
Torosaurus ("Perforated lizard")
was a genus of ceratopsid dinosaur. It had one of the largest
skulls of any land animal known, reaching 2.6 meters (8.5 ft)
in length. From head to tail, Torosaurus probably measured
about 7.6 meters (25 ft) long and weighed an estimated 4 to 6
tonnes (4.4 to 6.6 tons).
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Triceratops |
Triceratops is a genus of herbivorous
ceratopsid dinosaur which lived during the late Maastrichtian
stage of the Late Cretaceous Period, around 68 to 65 million years
ago (mya) in what is now North America. It was one of the last
dinosaur genera to appear before the great Cretaceous–Tertiary
extinction event. Individual Triceratops are estimated
to have reached about 7.9 to 9.0 m (26.0–29.5 ft) in length,
2.9 to 3.0 m (9.5–9.8 ft) in height, and 6.1–12.0
tonnes (13,000-26,000 lb) in weight. The most distinctive feature
is their large skull, among the largest of all land animals. It
could grow to be over 2 m (7 ft) in length, and could reach almost
a third of the length of the entire animal. It bore a single horn
on the snout, above the nostrils, and a pair of horns approximately
1 m (3 ft) long, with one above each eye. To the rear of the skull
was a relatively short, bony frill. Most other frilled dinosaurs
had large fenestrae in their frills, while the frills of Triceratops
were noticeably solid.
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